Structure Learning and Causal Learning

Boorman, E.D., Rajendran, V., O’Reilly, J.X., Behrens, T.E. (2016). Two computationally and anatomically distinct learning signals predict changes to stimulus-outcome associations in hippocampus. Neuron: 89:1343-54. article

Boorman, E.D., Rushworth, M.F. (2009). Conceptual representation and the making of new decisions. Neuron, 63:721-3. article


Domain Generality: Reward-based and Social Learning

Hill, M.R., Boorman, E.D., Fried, I. (2016). Observational learning computations in single neurons of the human anterior cingulate cortex. Nature Communications: 7:12722 article

Boorman, E.D., O’Doherty, J.P., Adolphs, R., Rangel, A. (2013). The behavioral and neural mechanisms underlying the tracking of expertise. Neuron: 80:1558-71. article


Behavioral Adaptation

Boorman, E.D., Rushworth, M.F., Behrens, T.E. (2013). Ventromedial prefrontal and anterior cingulate cortex adopt choice and default reference frames during sequential multialternative choice. Journal of Neuroscience: 33:2242-53. article

Mobbs, D. Hassabis, D., Yu, R., Chu, C., Rushworth, M., Boorman, E., Dalgleish, T. (2013). Foraging under competition: The neural basis of input matching in humans. Journal of Neuroscience: 33: 9866-72. article

Boorman, E.D., Behrens, T.E., Rushworth, M.F. (2011). Counterfactual choice and learning in a neural network centered on human lateral frontopolar cortex. PLoS Biology: 9:e1001093. article

Boorman, E.D., Behrens, T.E., Woolrich, M.W., and Rushworth, M.F.S. (2009). How green is the grass on the other side? Frontopolar cortex and the representation of alternative courses of action. Neuron, 62:733-43. article


Valuation and Choice

Croxson, P., Walton, M.E., Boorman, E.D., Rushworth, M.F., Bannerman, D.M. (2014). Unilateral medial frontal cortex lesions cause a cognitive decision-making deficit in rats. Eur J Neurosci. 40: 3757-65. article

Rushworth, M.F., Noonan, M.P., Boorman, E.D., Walton, M.E., Behrens, T.E. (2011). Frontal cortex and reward-guided learning and decision-making. Neuron: 70:1054:69. 

Boorman, E.D., Sallet, J. (2009). Mean-variance or prospect theory? The nature of value representations in the human brain. Journal of Neuroscience. 29:7945-7. 

Boorman, E.D., O’Shea, J., Sebastian, C., Rushworth, M.F.S., Johansen-Berg, H. (2007). Individual Differences in White-Matter Microstructure Reflect Variation in Functional Connectivity during Choice, Current Biology, 17:1426-31. article


Action Control

Higo, T., Mars, R.B., Boorman, E.D., Buch, E.R., Rushworth, M.F. (2011). Distributed and causal influence of frontal operculum in task control. PNAS: 108:4230-5. article

Buch, E.R., Mars, R.B., Boorman, E.D., Rushworth, M.F. (2010). A network centered on ventral premotor cortex exerts both facilitatory and inhibitory control over primary motor cortex during action reprogramming. Journal of Neuroscience. 30:1395-401. article

Mars, R.B., Klein, M.C., Neubert, F.X., Olivier, E., Buch, E.R., Boorman, E.D., Rushworth, M.F. (2009). Short-latency influence of medial frontal cortex over primary motor cortex during action selection under conflict. Journal of Neuroscience, 29:6926-31. article

Siebner HR, Bergmann TO, Bestmann S, Massimini M, Johansen-Berg H, Mochizuki H, Bohning DE, Boorman ED, et al. Consensus paper: combining transcranial stimulation with neuroimaging. (2009). Brain Stimulation. 2:58-80. article

O’Shea, J., Sebastian, C., Boorman, E.D., Johansen-Berg, H., and Rushworth, M.F.S. (2007). Functional specificity of premotor-motor cortical interactions during action selection. European Journal of Neuroscience, 26:2085-95. article



Bhagwagar Z, Wylezinska M, Jezzard P, Evans J, Boorman E, Matthews PM, Cowen P J (2008). Low GABA concentrations in occipital cortex and anterior cingulate cortex in medication-free, recovered depressed patients. Int J Neuropsychopharmacol., 11:255-60. article


Book chapters

Jocham, G, Boorman, ED, Behrens, TE (2015). The neurobiology of value-based decision making. In: Robin Murphy and Rob Honey (Eds.) The Wiley handbook on the Cognitive Neuroscience of learning. Wiley-Blackwell. 

Boorman, ED and Noonan, MP (2011). Contributions of ventromedial prefrontal and frontal polar cortex to reinforcement learning and value-based choice. In: Rogier Mars, Jerome Sallet, Matthew Rushworth, Nick Yeung (Eds.) Neural basis of motivational and cognitive control. Cambridge: MIT Press.

Rushworth, MFS, Boorman, E, Mars, RB (2009). Comparing brain connections in different species using diffusion-weighted imaging. In: Johansen-Berg H & Behrens TEJ (Eds.) Imaging brain pathways - Diffusion MRI: From quantitative measurement to in-vivo neuroanatomy. Amsterdam: Elsevier.